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Holman, A. J., & Fritz, U. (2001). A new emydine species from the middle miocene (barstovian) of nebraska, usa with a new generic arrangement for the species of clemmys sensu mcdowell (1964). Zoologische Abhandlungen / Staatliches Museum für Tierkunde in Dresden, 51(2), 331–353. 
Added by: Admin (17 Aug 2008 18:17:18 UTC)   Last edited by: Beate Pfau (08 Nov 2008 18:57:18 UTC)
Resource type: Journal Article
BibTeX citation key: Holman2001a
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Categories: General
Keywords: Actinemys, Actinemys marmorata, Clemmys, Clemmys guttata, Emydidae, Glyptemys, Glyptemys insculpta, Glyptemys muhlenbergii, Schildkröten = turtles + tortoises, Systematik = taxonomy, Testudinidae, Testudo
Creators: Fritz, Holman
Collection: Zoologische Abhandlungen / Staatliches Museum für Tierkunde in Dresden
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Abstract     
Testudinidae Abstract: As there is convincing evidence for a paraphyly of the genus Clemmys sensu McDOWELL (1964), it is suggested that Clemmys RITGEN, 1828 be restricted to its type species, Clemmys guttata (SCHNEIDER, 1792). Clemmys marmorata (BAIRD & GIRARD, 1852), a species more closely related to emydine taxa with a hinged plastron than to other species with nonhinged plastra previously referred to Clemmys, is treated as a member of the monotypic genus Actinemys AGASSIZ, 1857. Clemmys insculpta (LE CONTE, 1830 (1829)) and Clemmys muhlenbergii (SCHOEPFF, 1801) are regarded as representing another distinct genus for which the name Glyptemys AGASSIZ, 1857 (species typica: Testudo insculpta LE CONTE, 1830 (1829)=Glyptemys insculpta) is given precedence over Calemys AGASSIZ, 1857, erected for Testudo muhlenbergii SCHOEPFF, 1801. - Based on a large amount of fossil material from Nebraska (USA), Glyptemys valentinensis n. sp. is described. With this new species, the genus Glyptemys is now known from the Middle Miocene (Medial to Late Barstovian: ca 14.5-11.5 Ma BP) to modern times. Glyptemys valentinensis is morphologically more similar to Glyptemys insculpta than to G. muhlenbergii. It is suggested that G. valentinensis gave rise to G. insculpta between Late Barstovian and Late Hemphillian times (11.5-5.5 Ma BP). As sequence data of the mitochondrial cytochrome b gene argue for a differentiation of G. insculpta and G. muhlenbergii exactly in Medial Barstovian times (ca 14.5 Ma BP), G. valentinensis could be the last common ancestor of G. insculpta and G. muhlenbergii.
Added by: Admin  Last edited by: Beate Pfau
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