| Arnold, N. E. (1979). Indian ocean giant tortoises: their systematics and island adaptations. Philosophical Transactions of the Royal Society of London. Series B, 286(1011), Biological. |
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| Austin, J. J., Arnold, N. E., & Bour, R. (2003). Was there a second adaptive radiation of giant tortoises in the indian ocean? using mitochondrial dna to investigate speciation and biogeography of aldabrachelys (reptilia, testudinidae). Molecular Ecology, 12(6), 1415–1424. |
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| Burleigh, R., & Arnold, N. E. (1986). Age and dietary differences of recently extinct indian ocean tortoises (geochelone s. lat.) revealed by carbon isotope analysis. Proceedings of the Royal Society of London - Biological Sciences, 227(1246), Biological. |
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